Dusky grouper (Epinephelus marginatus)

General data

Main name: Dusky grouper
Local names: Yellowbelly rock cod, Yellowbelly grouper
Climates: Tropical, Subtropical, Temperate
Habitat: Saltwater
Native: Europe, Africa, Asia, South America
Distribution: Mediterranean Sea, Atlantic Ocean, Indian ocean

Classification

Description

Epinephelus marginatus, the dusky grouper, yellowbelly rock cod or yellowbelly grouper, is a species of marine ray-finned fish, a grouper from the subfamily Epinephelinae which is part of the family Serranidae, which also includes the anthias and sea basses. This species is the best-known grouper species of the Mediterranean Sea and North Africa coast. Description Epinephelus marginatus is a very large, oval-bodied and large-headed fish with a wide mouth which has a protruding lower jaw. The head and upper body are coloured dark reddish-brown or greyish, usually with yellowish gold countershading on the ventral surfaces; the base colour is marked by a vertical series of irregular pale greenish-yellow or silvery grey or whitish blotching which is normally rather conspicuous on the body and head; the black maxillary streak varies in its markedness; dark brown median fins; distal edges of the anal and caudal fins and also often pectoral fins have narrow white terminal bands; the pelvic fins are black towards their tips while the pectoral fins are dark reddish-brown or grey; the margin of the spiny dorsal fin and basal part of the pectoral fins are often golden yellows in colour. There are eleven spines and 13-16 soft rays in the dorsal fin. This species can grow up to 150 cm in standard length but is more often 90 cm. Distribution Epinephelus marginatus has two disjunct distribution centres, the main one is in the eastern Atlantic from the west coast of Iberia south along the western coast of Africa to the Cape of Good Hope, extending east into the south-western Indian Ocean, as far as southern Mozambique, with doubtful records from Madagascar and possibly Oman. It is found throughout the Mediterranean too. The second population occurs in the southwestern Atlantic off the coast of South America in southern Brazil, Uruguay and northern Argentina. In the eastern Atlantic it is not normally found further north than Portugal but there have been rare records from the Bay of Biscay and in the English Channel as far north as northern France, Great Britain and Ireland. Habitat Epinephelus marginatus is demersal, normally found in and around rocky reefs from surface waters down to as much as 300 metres in depth. It often occurs in the vicinity of beds of Poseidonia seagrass. Juveniles are generally found more inshore than adult fish, even being found in rock pools. Where they are protected, in marine nature reserves and no-take zones, both adults and juveniles occur in shallow waters, but the depths at which juveniles are found is always shallower than the preferred depths of adults.

Taxonomy of Fishes

Osteichthyes - Bony fishes->
- Coelacanthi - Lobe-finned fishes->
- Teleostei - Ray-finned fishes->
  - Perciformes - Perches->
    
    Percidae - Perches->
    
    Pomatomidae - Bluefishes->
    
    Serranidae - Sea basses->
    
    Pomacentridae - Damsel and clownfishes->
    
    Latidae - Lates perches->
    
    Centropomidae - Snooks->
    
    Lutjanidae - Snappers->
    
    Haemulidae - Grunts->
    
    Artedidraconidae - Barbled plunderfishes->
    
    Embiotocidae - Surfperches->
    
    Glaucosomatidae - Pearl perches->
    
    Malacanthidae - Tilefishes->
    
    Polynemidae - Threadfin->
    
    Gerreidae - Mojarras->
    
    Priacanthidae - Bigeyes->
    
    Monodactylidae - Moonyfishes->
    
    Centrogenyidae - False scorpionfishes->
    
    Epinephelidae - Groupers->
    
    Epinephelus - Cloudy groupers->
    
    Atlantic goliath grouper (Epinephelus itajara)
    
    Dusky grouper (Epinephelus marginatus)
    
    Goldblotch grouper (Epinephelus costae)
    
    Red grouper (Epinephelus morio)
    
    Nassau grouper (Epinephelus striatus)
    
    Malabar grouper (Epinephelus malabaricus)
    
    Maori rockcod (Epinephelus undulatostriatus)
    
    Blacktip grouper (Epinephelus fasciatus)
    
    One-blotch grouper (Epinephelus melanostigma)
    
    Comet grouper (Epinephelus morrhua)
    
    Pacific goliath grouper (Epinephelus quinquefasciatus)
    
    Giant grouper (Epinephelus lanceolatus)
    
    Honeycomb grouper (Epinephelus merra)
    
    Potato grouper (Epinephelus tukula)
    
    Blue-and-yellow grouper (Epinephelus flavocaeruleus)
    
    White grouper (Epinephelus aeneus)
    
    Epaulet grouper (Epinephelus stoliczkae)
    
    Rock hind (Epinephelus adscensionis)
    
    Hong Kong grouper (Epinephelus akaara)
    
    White-edged grouper (Epinephelus albomarginatus)
    
    Banded grouper (Epinephelus amblycephalus)
    
    Spotted grouper (Epinephelus analogus)
    
    Catface grouper (Epinephelus andersoni)
    
    Areolate grouper (Epinephelus areolatus)
    
    Yellow grouper (Epinephelus awoara)
    
    Frostback rockcod (Epinephelus bilobatus)
    
    Duskytail grouper (Epinephelus bleekeri)
    
    Palemargin grouper (Epinephelus bontoides)
    
    Longtooth grouper (Epinephelus bruneus)
    
    Dogtooth grouper (Epinephelus caninus)
    
    Moustache grouper (Epinephelus chabaudi)
    
    Tonga grouper (Epinephelus chlorocephalus)
    
    Brownspotted grouper (Epinephelus chlorostigma)
    
    Epinephelus clippertonensis (Epinephelus clippertonensis)
    
    Whitespotted grouper (Epinephelus coeruleopunctatus)
    
    Orange-spotted grouper (Epinephelus coioides)
    
    Coral grouper (Epinephelus corallicola)
    
    Brokenbar Grouper (Epinephelus craigi)
    
    Speckled blue grouper (Epinephelus cyanopodus)
    
    Saddletail grouper (Epinephelus daemelii)
    
    Spinycheek grouper (Epinephelus diacanthus)
    
    Speckled hind (Epinephelus drummondhayi)
    
    Dotted grouper (Epinephelus epistictus)
    
    Cloudy grouper (Epinephelus erythrurus)
    
    Rock grouper (Epinephelus fasciatomaculosus)
    
    Barred-chest grouper (Epinephelus faveatus)
    
    Brown-marbled grouper (Epinephelus fuscoguttatus)
    
    Darkmargin grouper (Epinephelus fuscomarginatus)
    
    Multispotted grouper (Epinephelus gabriellae)
    
    Epinephelus geoffroyi (Epinephelus geoffroyi)
    
    Dungat grouper (Epinephelus goreensis)
    
    Red hind (Epinephelus guttatus)
    
    Bridled grouper (Epinephelus heniochus)
    
    Starspotted grouper (Epinephelus hexagonatus)
    
    Blacksaddle grouper (Epinephelus howlandi)
    
    Somali grouper (Epinephelus indistinctus)
    
    Pacifc insular grouper; (Epinephelus insularis)
    
    Marquesan grouper (Epinephelus irroratus)
    
    Epinephelus japonicus (Epinephelus japonicus)
    
    Fivebar grouper (Epinephelus kupangensis)
    
    Starry grouper (Epinephelus labriformis)
    
    Striped grouper (Epinephelus latifasciatus)
    
    Mystery grouper (Epinephelus lebretonianus)
    
    Longspine grouper (Epinephelus longispinis)
    
    Snubnose grouper (Epinephelus macrospilos)
    
    Highfin grouper (Epinephelus maculatus)
    
    Speckled grouper (Epinephelus magniscuttis)
    
    Netfin grouper (Epinephelus miliaris)
    
    White-blotched grouper (Epinephelus multinotatus)
    
    White-streaked grouper (Epinephelus ongus)
    
    Dot-dash grouper (Epinephelus poecilonotus)
    
    Smallscaled grouper (Epinephelus polylepis)
    
    Camouflage grouper (Epinephelus polyphekadion)
    
    White-dotted grouper (Epinephelus polystigma)
    
    Striped-fin grouper (Epinephelus posteli)
    
    Longfin grouper (Epinephelus quoyanus)
    
    Oblique-banded grouper (Epinephelus radiatus)
    
    Red-tipped grouper (Epinephelus retouti)
    
    Halfmoon grouper (Epinephelus rivulatus)
    
    Sixbar grouper (Epinephelus sexfasciatus)
    
    Surge grouper (Epinephelus socialis)
    
    Foursaddle grouper (Epinephelus spilotoceps)
    
    Black-dotted grouper (Epinephelus stictus)
    
    Seamount grouper (Epinephelus suborbitalis)
    
    Summan grouper (Epinephelus summana)
    
    Greasy grouper (Epinephelus tauvina)
    
    Yellowspotted grouper (Epinephelus timorensis)
    
    Threespot grouper (Epinephelus trimaculatus)
    
    Plump grouper (Epinephelus trophis)
    
    Reticulate grouper (Epinephelus tuamotuensis)
    
    Wavy-lined grouper (Epinephelus undulosus)
    
    Mycteroperca - Groupers ->
    
    Cephalopholis - Scaleheaded groupers->
    
    Cromileptes - Humpback groupers->
    
    Hyporthodus - Hyporthodus->
    
    Alphestes - Alphestes->
    
    Variola - Variola->
    
    Anyperodon - Anyperodon->
    
    Gonioplectrus - Gonioplectrus->
    
    Plectropomus - Coral groupers->
    
    Gracila - Gracila->
    
    Aethaloperca - Aethaloperca->
    
    Chromileptes - Chromileptes->
    
    Dermatolepis - Dermatolepis->
    
    Saloptia - Saloptia->
    
    Triso - Triso->
    
    Cebidichthyidae - Cebidichthyidae->
    
    Anthiadidae - Anthiadidae->
    
    Ambassidae - Asiatic glassfishes->
    
    Bathydraconidae - Antarctic dragonfishes->
    
    Bembropidae - Duckbill flatheads->
    
    Bovichtidae - Thornfishes->
    
    Caesionidae - Fusiliers->
    
    Caesioscorpididae - Blowhole perches->
    
    Centracanthidae - Picarels->
    
    Cepolidae - Bandfishes->
    
    Channichthyidae - Crocodile icefishes->
    
    Congrogadidae - Eelblennies->
    
    Datnioididae - Freshwater tripletails->
    
    Dinopercidae - Cavebasses->
    
    Eleginopidae - Patagonian blennies->
    
    Grammatidae - Basslets->
    
    Grammistidae - Soapfishes->
    
    Hapalogenyidae - Barbeled grunters->
    
    Harpagiferidae - Spiny plunderfishes->
    
    Lactariidae - False trevallies->
    
    Latilidae - Deepwater tilefishes->
    
    Leptobramidae - Beachsalmon->
    
    Liopropomatidae - Painted basslets->
    
    Niphonidae - Ara groupers->
    
    Nototheniidae - Cod icefishes->
    
    Opistognathidae - Jawfishes->
    
    Oxylebiinae - Painted greenlings->
    
    Plesiopidae - Roundheads->
    
    Pseudaphritidae - Catadromous icefishes->
    
    Pseudochromidae - Dottybacks->
    
    Ptereleotridae - Dartfishes->
    
    Toxotidae - Archerfishes->
  - Cypriniformes - Carps->
  - Esociformes - Pikes->
  - Salmoniformes - Salmons and Trouts->
  - Siluriformes - Catfishes->
  - Scorpaeniformes - Mail-cheeked fishes->
  - Gasterosteiformes - Sticklebacks->
  - Gadiformes - Cods->
  - Anguilliformes - Eels and morays->
  - Clupeiformes - Herrings->
  - Syngnathiformes - Pipefishes and Seahorses->
  - Gobiiformes - Gobies->
  - Mugiliformes - Mullets->
  - Carangiformes - Jacks->
  - Beloniformes - Needlefishes->
  - Labriformes - Wrasses->
  - Scombriformes - Mackerels->
  - Mulliformes - Goatfishes->
  - Pleuronectiformes - Flatfishes->
  - Aulopiformes - Grinners->
  - Blenniiformes - Blennies->
  - Cyprinodontiformes - Toothcarps->
  - Osmeriformes - Smelts->
  - Centrarchiformes - Basses and sunfishes->
  - Cichliformes - Cichlids->
  - Characiformes - Characins->
  - Osteoglossiformes - Bony tongues->
  - Elopiformes - Tarpons and tenpounders->
  - Anabantiformes - Gouramies and snakeheads->
  - Gymnotiformes - South American knifefish->
  - Istiophoriformes - Barracudas->
  - Albuliformes - Bonefishes->
  - Lepidogalaxiiformes - Salamanderfishes->
  - Lophiiformes - Anglerfishes->
  - Spariformes - Breams and porgies->
  - Trachiniformes - Weeverfishes->
  - Moroniformes - Temperate basses->
  - Acanthuriformes - Surgeonfishes->
  - Zeiformes - Dories->
  - Tetraodontiformes - Puffers and filefishes->
  - Holocentriformes - Squirrelfishes->
  - Dactylopteriformes - Flying gurnards->
  - Lampriformes - Lamprids->
  - Galaxiiformes - Southern smelts->
  - Batrachoidiformes - Toadfishes->
  - Hiodontiformes - Mooneyes->
  - Percopsiformes - Trout-perches->
  - Callionymiformes - Dragonets->
  - Atheriniformes - Silversides->
  - Notacanthiformes - Spiny eels->
  - Saccopharyngiformes - Swallowers and Gulpers->
  - Alepocephaliformes - Slickheads and tubeshoulders->
  - Gonorynchiformes - Milkfishes->
  - Argentiniformes - Marine smelts->
  - Stomiiformes - Lightfishes and dragonfishes->
  - Ateleopodiformes - Jellynose fishes->
  - Myctophiformes - Lanternfishes->
  - Stylephoriformes - Tube-eyes->
  - Polymixiiformes - Beardfishes->
  - Beryciformes - Sawbellies->
  - Trachichthyiformes - Rughies->
  - Ophidiiformes - Cusk-eels->
  - Kurtiformes - Nurseryfishes & Cardinalfishes->
  - Synbranchiformes - Swamp eels->
  - Gobiesociformes - Clingfishes->
  - Acropomatiformes - Oceanic basses->
- Cladistii - Bichirs->
- Dipneusti - Lungfishes->
- Chondrostei - Sturgeons->
- Holostei - Gars and Bowfins->
Chondrichthyes - Cartilaginous fishes->
Agnatha - Jawless fishes->